People
Desmond R. Layne
Professor of Pomology
Endowed Chair – Tree Fruit Extension Program Leader
(WSU faculty member since 2013)
Areas of Interest
- Germplasm evaluation and new cultivar development
- Orchard systems management
- Innovative tools for extension outreach
- International training programs for fruit growers
Educational Background
- Ph.D., Horticulture (pomology emphasis), Michigan State University
- M.S., Horticulture (pomology emphasis), Michigan State University
- B.Sc., Agriculture (horticulture major), University of Guelph, Ontario, Canada
Work Experience
- 1997-2012: Professor of Pomology, State Fruit Specialist and State Extension Program Team Leader (Horticulture), Clemson University, Clemson, SC
- 1993-1997: Principal Investigator of Horticulture, Kentucky State University, Frankfort, KY
- 1992-1993: Research Plant Physiologist, USDA Forestry Sciences Laboratory, Rhinelander, WI
Selected Publications
Layne, D.R., Z. Wang and L. Niu. 2013. Protected cultivation of peach and nectarine in China – Industry observations and assessments. J. Amer. Pom. Sci. 67(1):18-28.
Sparks, B., J. Frecon, and D.R. Layne. 2012. Mr. Peach and The Peach Doctor – two respected peach veterans weigh in on varieties, how to increase consumption and more. The American Fruit Grower magazine 132(7): cover and p. 8-11.
Layne, D.R. and D. Bassi (eds). 2008. The Peach: Botany, Production and Uses. 2008. CAB International Press, Wallingford, U.K. 615 pp.
Okie, W.R. and D.R. Layne. 2008. Early Augustprince and Augustprince peaches. HortScience 43(5):1600-1602. (new peach cultivar releases)
Okie, W.R. and D.R. Layne. 2008. Scarletprince and Julyprince peaches. HortScience 43(5):1603-1605. (new peach cultivar releases)
Pomper, K.W. and D.R. Layne. 2005. The North American pawpaw: botany and horticulture. Horticultural Reviews 31:351-384.
Schnabel, G., and D.R. Layne. 2004. Comparison of reduced-application and sulfur-based fungicide programs on scab intensity, fruit quality, and cost of disease control of peach. Plant Disease 88:162-166.
Layne, D.R., D.B. Cox, and E.J. Hitzler. 2002. Peach systems trial: the influence of training system, tree density, rootstock, irrigation, and fertility on growth and yield of young trees in South Carolina. Acta Hort. 592(1):367-376.
Layne, D.R., Z. Jiang, and J.W. Rushing. 2002. The influence of ReTain and reflective film on color and maturity of ‘Gala’ apples. HortTechnology 12(4):640-645.
Layne, D.R., Z. Jiang, and J.W. Rushing. 2001. Reflective film improves red skin coloration and advances maturity in peach. HortTechnology 11(2):234-242.
Rita L. Hummel
Associate Professor/Associate Scientist/Extension Horticulturist
(WSU faculty member since 1985)
Areas of Interest
Environmental Horticulture
- Develop recycled organic materials as substrates and nutrient sources for nursery and greenhouse container production and landscape use
- Enhance water conservation in container production and landscapes
- Improve longevity of trees and shrubs in the landscape by preventing root structural defects
- Develop production practices for difficult to grow native plants
- Determine woody plant adaptation to rain gardens under Pacific Northwest environmental conditions
Educational Background
- Ph.D., University of Minnesota, Horticulture
- M.S., University of Minnesota, Horticulture
- B.S., South Dakota State University, Botany, Horticulture
Awards/Memberships
- Washington State Nursery and Landscape Association Educator of the Year 2009
- USDA Plant Hardiness Zone Map Technical Review Team
Teaching Responsibilities
Program Personnel
- Robert Riley, Professional Worker II
- Ian Grinter, MS (2008-2012)
Selected Publications
- Hummel, R.L. and C. Cogger. 2012. Struvite can serve as a renewable phosphorus source for greenhouse crop production. B&B Magazine 64(4):16-17.
- CSANR Webinar ‘Struvite as a Phosphorus Fertilizer Source for Greenhouse Crop Production.’ http://breeze.wsu.edu/p29658863/
- Ophardt, M.C., and R. L. Hummel. 2011. Planting Trees and Shrubs in the Landscape. WSU Ext. Fact Sheet FS047E. pp 10. http://cru.cahe.wsu.edu/CEPublications/FS047E/FS047E.pdf
- Krucker, M., R.L. Hummel, and C. Cogger. 2010. Chrysanthemum production in composted and noncomposted organic waste substrates fertilized with nitrogen at two rates using surface and subirrigation. HortScience. 45(11):1695-1701.
- Scagel, C., R. Regan, R. L. Hummel, and G. Bi. 2010. Cold tolerance of container-grown green ash trees is influenced by nitrogen fertilizer type and rate. HortTechnology. 20(2):292-303.
- Hummel, R.L., E. Evans, and R. Riley. 2009. Effect of Mechanical Root Disruption at Transplant on Short- and Longer-term Growth and Survival of Scotch Pine (Pinus sylvestris) and Shore Pine (Pinus contorta var. contorta). pp 211-221. In: G.W. Watson, L. Costello, B. Scharenbroch and E. Gilman (eds.), The Landscape Below Ground III: Proceedings of an International Workshop on Tree Root Development in Urban Soils. International Society of Arboriculture, The Morton Arboretum, Lisle, IL.
- Cogger, C., R. Hummel, J. Hart, and A. Bary. 2008. Soil and redosier dogwood response to incorporated and surface-applied compost. HortScience. 43(7):2143-2150. Read more »
Linda Chalker-Scott
Associate Professor and Extension Specialist
(WSU faculty member since 2004)
For a complete profile, click here.
Area of Interest
- Urban horticulture – woody landscape plants, restoration horticulture
Educational Background
- Ph.D., Horticulture (minors in Biochemistry & Botany), Oregon State University (OSU)
- M.S., Marine Biology, OSU
- B.S., General Biology, OSU
Teaching Responsibilities
- Guest lectures, WSU and University of Washington
Program Personnel
- Irena Neffeova, MS (2011-2013)
Selected Publications
Chalker-Scott, L. 2012. Protecting water resources: planting and caring for home wetlands. WSU Extension Fact Sheet.
Gillman, J., L. Chalker-Scott, H. Scoggins and B. Cregg. 2011. A successful multi-institutional blog for transferring garden and landscape information to the public. Journal of Extension. Available online at: http://www.joe.org/joe/2011december/iw3.php.
Chalker-Scott, L. 2010. The Informed Gardener Blooms Again. University of Washington Press, Seattle, WA.
Chalker-Scott, L. 2009. Sustainable Landscapes and Gardens: good science – practical application. GFG Publishing, Yakima, WA.
Chalker-Scott, L. 2008. The Informed Gardener. University of Washington Press, Seattle, WA.
Chalker-Scott, L. 2007. Impact of mulches on landscape plants and the environment – a review. Journal of Environmental Horticulture 25(4): 239-249.
Chalker-Scott, L. 2002. Do anthocyanins function as osmoregulators in leaf tissues? Advances in Botanical Research 37:103-127.
Reichard, S., L. Chalker-Scott and S. Buchanan. 2001. Interactions among non-native plants and birds, pp. 179-223. In: J. Marzluff, R.A. Bowman, and R. Donnelly (eds.), Avian Ecology and Conservation in an Urbanizing World, Kluwer Academic Publishers, Boston, MA.
Chalker-Scott, L. 1999. Environmental significance of anthocyanins in plant stress responses. Invited review, Photochemistry and Photobiology 70:1-9.
Bhaskar Bondada
Assistant Professor
(WSU faculty member since 2004)
For a complete profile, click here.
Areas of interest
- Grapes; microscopy and physiology of grape and vine
Educational Background
- Ph.D., University of Arkansas, Fayetteville, AR
- M.S., Tamil Nadu Agricultural University, Coimbatore, India
- B.S., Tamil Nadu Agricultural University, Coimbatore, India
Teaching Responsibilities
- Advanced Viticulture
- Other courses- Small Fruits and Viticulture, Seminars in Viticulture and Enology, Wine making
Current Graduate Students
- Susan Hill
Selected Publications
RESEARCH
Bondada, B. and J. Shutthanandan. 2012. Physio-anatomical responses of leaf and fruit of grapevine (Vitis vinifera) to re-watering following water stress. American Journal of Plant Science (In Review)
Keller, M., P. M. Shrestha, M. Biondi, and B. Bondada. 2012. Phloem inflow and apoplastic unloading in ripening grape berries drive xylem backflow. Plant Physiology (In Review).
Bondada, B. and M. Keller. 2012. Not all shrivels are created equal – morpho-anatomical and compositional characteristics vary among different shrivel forms that develop during ripening of grape (Vitis vinifera L.) berries. American Journal of Plant Science (In Press).
Bondada, B. and M. Keller. 2012. Morpho-anatomical symptomatology and osmotic behavior of grape berry shrivel. Journal of the American Society for Horticultural Science 137:20-30.
Bondada, B. 2012. Technical Advance: Novel, Simple, Fast, and Safe Approaches to Visualizing Fine Cellular Structures in Free-hand Sections of Stem, Leaf, and Fruit Using Optical Microscopy. Current Botany 3:11-22.
Bondada, B. 2011. Anomalies in structure, growth characteristics, and nutritional composition as induced by 2, 4-D drift phytotoxicity in grapevine (Vitis vinifera L.) leaves and clusters. Journal of the American Society for Horticultural Science 136:165-176.
Bondada, B. 2011. Micromorpho-anatomical examination of 2, 4-D phytotoxicity in grapevine (Vitis vinifera L.) leaves. Journal of Plant Growth Regulation 30:185-198.
EXTENSION
Bondada, B. 2012. Ripening disorders of Grape. Grape IPM Manual.
Graduate Students
Eliane Thaines BodahDegree sought: Ph.D. Areas of interest: Sustainable food production, molecular breeding, agricultural ecosystems Educational background: M.S., Education, University of Passo Fundo, Brazil; B.S., Biology, Federal University of Santa Maria, Brazil Faculty advisor: Amit Dhingra |
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Jeremy CowanDegree sought: Ph.D. Areas of interest: Fruit and vegetable horticulture, sustainable production Educational background: M.B.A., Marketing and New Venture Management, University of San Diego; B.S., Horticulture, Brigham Young University Faculty advisor: Carol Miles |
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James CrabbDegree sought: Ph.D. Area of interest: Wine grape genomics Educational background: B.S., Horticulture, WSU Faculty advisor: Amit Dhingra |
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Rachel DanielDegree sought: Ph.D. (Molecular Plant Science) Areas of interest: Genetics and genomics, apples Educational background: B.S., Integrated Plant Sciences, Agricultural Biotechnology, WSU. Faculty advisor: Amit Dhingra |
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Luke GustafsonDegree sought: M.S. Areas of interest: Sustainable vegetable production, plant-insect interactions Educational background: B.S., Biology; B.A., Spanish, University of Minnesota, Duluth Faculty advisor: Preston Andrews |
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Matthew HalldorsonArea of interest: Viticulture Educational background: B.S., Plant Biology, Northern Arizona University Faculty advisor: Markus Keller |
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Julia HarshmanDegree sought: Ph.D. Area of interest: Plant breeding, tree architecture, tree and fruit physiology Educational background: M.S., Plant Sciences, University of Maryland, College Park; B.S., Plant Sciences, University of Maryland, College Park Faculty advisor: Kate Evans |
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Christopher HendricksonAreas of interest: Genetic mechanisms in the regulation of climacteric fruit ripening, and the role of chilling in ripening induction Educational background: M.S., Plant Physiology, Utah State University; B.S., Plant Science Research, Rutgers University, NJ Faculty advisor: Amit Dhingra |
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Derek HermanDegree sought: M.S. Area of interest: Post Harvest Physiology of Small Fruit and Vegetables, Potato Agronomy Educational background: B.S. Horticulture, emphasis Viticulture and Enology, WSU Faculty advisor: Rick Knowles |
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Sacha JohnsonArea of interest: Vegetable horticulture Educational background: B.S., Ecosystem Science and Resource Management, University of Washington Faculty advisor: Carol Miles |
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Brittany KommDegree sought: M.S. Areas of interest: Viticulture and canopy management Educational background: B.S., Horticulture – Fruit and Vegetable Management, Viticulture emphasis, WSU Faculty advisor: Michelle Moyer |
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Yu MaDegree sought: Ph.D. Areas of interest: Pea genomic assisted breeding and bioinformatics Educational background: M.S., Crop Germplasm Resources, Chinese Academy of Agricultural Sciences, China; B.S., Biotechnology, Agricultural University of Hebei, China Faculty advisor: Dorrie Main |
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Irena NeffeovaDegree sought: M.S. Areas of interest: Root development in woody plants after transplanting, WSU Women’s Rowing Educational background: B.S., Environmental Horticulture, WSU Faculty advisors: Virginia Lohr and Linda Chalker-Scott |
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Sushan RuDegree sought: Ph.D. Area of interest: Bioinformatics and its application in plant breeding Educational background: M.S., Horticulture, WSU; B.S., Horticulture, Southwest University, China Faculty advisor: Dorrie Main |
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Rhett SpearDegree sought: Ph.D. Area of interest: Potato agronomy Educational background: B.S., Chemistry, Utah State University Faculty advisor: Mark Pavek |
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Sujeet VermaDegree sought: Ph.D. Areas of interest: Finding functionalhaplotypes for apple fresh sensation: higher acidity, higher crispness, and higher juiciness via pedigree-based QTL analysis and developing predictive genetic tests for fresh sensation; Plant physiology: pre- and post-harvest Educational background: M.S., Horticulture, WSU; B. Pharma, HNBG University, Uttaranchal, India. Faculty advisor – Cameron Peace |
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Xiaoyue (Katherine) WangDegree sought: M.S. Area of interest: Drought resistant cultivars of grapevines Educational background: B.S., Horticulture, China Agricultural University Faculty advisor: Bhaskar Bondada |
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Yun ZhangAreas of interest: Viticulture, plant water relations Educational background: B.S., Agricultural Water Resources Engineering, Northwest A&F University, Yangling, China Faculty advisor: Markus Keller |
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Yunyang ZhaoDegree sought: Ph.D. Area of interest:Understanding the genetics of pedicel fruit retention force in sweet cherry (Prunus avium L.) Educational background: M.S., Plant Genetics,Northeast Normal University, China; B.S., Biological Sciences, Northeast NormalUniversity, China Faculty advisor: Nnadozie Oraguzie |
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Daniel ZommickArea of interest: Biochemical pathways of cold-induced sweetening (CIS) in novel potato varieties Educational background: B.S., Horticulture – Environmental Horticulture, WSU Faculty advisor: Rick Knowles |
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Matthew D. Whiting
Associate Scientist/Associate Professor/Extension Specialist
(WSU faculty member since 2002)
For a complete profile, click here.
Areas of Interest
- Whole-tree physiology
- High efficiency orchard architecture
- Incorporating automation/mechanization in fruit production systems,
- Environmental control of fruit quality;
- Pollination biology
Educational Background
- Ph.D., Whole-tree physiology, Washington State University
- M.Sc., Postharvest physiology, University of Guelph
- B.Sc., Horticultural Science and Business, University of Guelph
Teaching Responsibilities
- HORT 503 – Critical Analysis of Scientific Literature (Fall 2011)
Program Personnel
- Lu Zhang (PhD, 2013)
- Yiannis Ampatzidis, Post-doctoral Research Associate
- David Gibeault, Post-doctoral Research Associate
Selected Publications
Click here to view.
Cameron P. Peace
Associate Scientist/Associate Professor
(WSU faculty member since 2006)
Areas of Interest
- DNA-informed breeding for tree fruit: turning promise into reality
- Co-Project Director for $14.4 M SCRI project RosBREED: Enabling Marker-Assisted Breeding in Rosaceae
- Team Leader for Marker-Assisted Breeding Pipeline and key member of Breeding Info. Manag. System, Pedigree-Based Analysis, Genomics, Genotyping, and Extension Teams
- Leader and founder of the Pacific Northwest Tree Fruit Genotyping Laboratory, (Pullman, WA)
Educational Background
- Ph.D., Horticultural Genetics, University of Queensland, Brisbane, Australia
- Bachelor of Horticultural Science, University of Queensland
Teaching Responsibilities
- HORT/CropS 202 – Crop Growth and Development
Program Personnel
- Daniel Edge-Garza, Associate in Research
- Terry Rowland, Genetic Screening Technician for the Pacific Northwest Tree Fruit Genotyping Laboratory
- Sujeet Verma (PhD, 2009–2013)
- Sushan Ru (PhD, 2012–2015)
Selected Publications
Chagné, D., …(18 other authors)…, and C. Peace. 2012. Genome-wide SNP detection, validation, and development of an 8K SNP array for apple. PLoS One 7(2):e31745.
Verde, I., …(21 other authors)…, and C. Peace. 2012. Development and evaluation of a 9K SNP array for peach by internationally coordinated SNP detection and validation in breeding germplasm. PLoS One 7(4):e35668.
Peace, C., …(12 other authors). 2012. Development and evaluation of a genome-wide 6K SNP array for diploid sweet cherry and tetraploid sour cherry. PLoS ONE (submitted).
B. W. (Joe) Poovaiah
WSU Regents Professor
(WSU faculty member since 1975)
For a complete profile, click here.
Areas of Interest
- Calcium/calmodulin-mediated signaling in plants
Educational Background
- PhD, Utah State University, Logan, UT
Teaching Responsibilities
- HORT 416/516 – Advanced Horticultural Crop Physiology
Program Personnel
- Liqun Du, Research Associate Professor
- Chenjia Shen, Visiting Assistant Professor
- Pratyush Routray (PhD candidate, 2007-2012)
- Lei Zhang (PhD candidate, 2007-2012)
Selected Publications
Qiu, Y., Xi, J., Du, L. and Poovaiah, B.W. 2012. The function of calreticulin in plant immunity: New discoveries for an old protein. Plant Signaling & Behavior 7:1-4.
Qiu, Y., Xi, J., Du, L., Suttle, J. and Poovaiah, B.W. 2012. Coupling calcium/calmodulin-mediated signaling and herbivore-induced plant response through calmodulin-binding transcription factor AtSR1/CAMTA3. Plant Mol. Biol. 79:89-99 (with journal cover page).
Xi, J., Qiu, Y., Du, L. and Poovaiah, B.W. 2012. Plant-specific trihelix transcription factor AtGT2L interacts with calcium/calmodulin and responds to cold and salt stresses. Plant Sci. 185-186:274-280.
Qiu, Y., Xi, J., Du, L., Roje, S. and Poovaiah, B.W. 2012. A dual regulatory role of Arabidopsis calreticulin 2 in plant innate immunity. Plant J. 69(3):489–500.
Yang, T., Ali, G.S., Yang, L, Du, L., Reddy, A.S.N., Poovaiah, B.W. 2010. Calcium/calmodulin-regulated Receptor-like Kinase CRLK1 interacts with MEKK1 in Plants. Plant Signaling and Behavior 5:991-994.
Yang, T, Chaudhuri, S, Yang, L, Du, L, Poovaiah, B.W. 2010. A calcium/calmodulin-regulated member of the receptor-like kinase family confers cold tolerance in plants. J. Biol. Chem. 285:7119-7126.
Du, L., Ali, G. S., Simons, K. A., Hou J., Yang, T., Reddy, A.S.N. and Poovaiah, B. W. 2009. Ca2+/calmodulin regulates salicylic-acid-mediated plant immunity. Nature 457:1154-1158.
Yang, T., Du, L. and Poovaiah, B.W. 2007. Concept of redesigning proteins by manipulating calcium/calmodulin-binding domains to engineer plants with altered traits. Functional Plant Biology 34:343-352.
Gleason, C., Chaudhuri, S., Yang, T., Munoz-Gutierrez, A., Poovaiah, B.W. and Oldroyd, G.E.D. 2006. Nodulation independent of rhizobia induced by a calcium-activated kinase lacking autoinhibition. Nature. 441:1149-1152.
Du, L. and Poovaiah, B. W. 2005. Calcium/calmodulin is critical for brassinosteroid biosynthesis and plant growth. Nature. 437:741-745.
Mark J. Pavek
Associate Professor/Potato Specialist
(WSU faculty member since 2004)
Areas of Interest
- Potato agronomy and variety development
For more information, visit the WSU Potato website.
Educational Background
- Ph.D., Horticulture – Potato Agronomy, Washington State University
- M.S., Plant Science – Weed Science, University of Idaho
- B.Sc., Ag. Econ. – Agribusiness, University of Idaho
- B.Sc., Plant Science – Crop Production, University of Idaho
Work Experience
- Field Research Manager, NatureMark Potatoes – Monsanto, PNW
- Commercial Potato Producer/Assistant Farm Manager – SSI Land and Cattle, Idaho
- Product Development Assistant – Syngenta, Idaho
- Commercial potato, sugarbeet, wheat, barley, corn, and alfalfa production experience
Nnadozie Oraguzie
Associate Professor/Scientist
(WSU faculty member since 2008)
Areas of Interest
- Plant breeding and genetics including quantitative genetics, association genetics, population genetics and molecular genetics. My focus is on sweet cherry but I can also research on peach, apple and pear.
Educational Background
- Ph.D., Plant breeding and Genetics, Lincoln University, New Zealand
- MSc., Plant breeding and Genetics, University of Ibadan, Nigeria
- BSc., Botany, University of Jos, Nigeria
Work Experience
- 1998-2008: Senior Scientist, HortResearch, New Zealand
- 2006: JSPS Invitation Fellow (for 6 months), National Institute of Fruit Tree Science, Tsukuba, Ibaraki, Japan
- 2001-2002: STA Fellow, National Institute of Fruit Tree Science, Morioka, Japan
- 1997-98: Post-Doc, Canterbury University, Christchurch, New Zealand
- 1988-89: Biology Teacher, Federal School of Art & Science, Ondo, Nigeria
Teaching Responsibilities
- HORT 503 – Critical Analysis of Scientific Literature (team taught with Drs. Whiting, Moyer, and Keller)
Program Personnel
- Blessing Athanson, Technician
- Sue Watkins, Technician
- Yunyang Zhao (2009-2013)
- Murali Bellamkonda (2010-2013)
- Josephine Mgbechi-Ezeri (2012-2014)
Selected Publications
BOOK
Oraguzie, N.C., E.H.A. Rikkerink, S. Gardiner, and N. de Silva (eds). 2007. Association mapping in plants, ISBN 9780387358444. Springer, New York.
BOOK CHAPTERS
Rikkerink, E.H.A., N.C. Oraguzie, and S.E. Gardiner. 2007. Prospects of association mapping in perennial horticultural crops, p. 249-269. In: N.C. Oraguzie, E.H.A. Rikkerink, S.E. Gardiner, and N. de Silva (eds.). Association mapping in plants. Springer, New York.
Oraguzie, N.C. and P.L. Wilcox. 2007. An overview of association mapping, p. 1-9. In: N.C. Oraguzie, E.H.A. Rikkerink, S.E. Gardiner, and N. de Silva (eds.). Association mapping in plants. Springer, New York.
Oraguzie, N.C., P.L. Wilcox, and E. Rikkerink. 2007. Linkage disequilibrium, p. 10-39. In: N.C. Oraguzie, E.H.A. Rikkerink, S.E. Gardiner, and N. de Silva (eds.). Association mapping in plants. Springer, New York.
Oraguzie, N.C., E. Rikkerink, S. Gardiner, V.G. Bus, A. Currie, R. Rusholme, and R. Volz. 2004. A review of developments in breeding techniques and gene technology tools for new cultivar development in apple. Recent Advances in Genetics and Breeding 1: 223-257.
JOURNAL ARTICLES
Oraguzie, N., P. Alspach, R. Volz, C. Whitworth, C. Ranatunga, R. Weskett, and R. Harker. 2009. Postharvest assessment of fruit quality parameters in apple using both instruments and an expert panel. Postharvest Biology and Technology (in press).
Gharaghani, A., Z. Zamani, A. Talaia, N. Oraguzie, R. Fattahi, H. Hajnajari, C. Wiedow, and S. Gardiner. 2009. Genetic identity and relationships of Iranian apple (Malus x domestica) cultivars and landraces, wild Malus species and representative old apple cultivars based on SSR marker analysis. Genetic Resources and Crop Evolution (in press).
Chagné, D., C.M. Carlisle, C. Blond, R.K. Volz, C.J. Whitworth, N.C. Oraguzie, R.N. Crowhurst, A.C. Allan, R.V. Espley, R.P. Hellens, and S.E. Gardiner. 2007. Mapping candidate genes for red flesh and foliage colour in apple. BMC Genomics 8:212.
Oraguzie, N.C., R.K. Volz, C.J. Whitworth, H.C.M. Bassett, A.J. Hall, S.E. Gardiner. 2007. Influence of Md-ACS1 allelotype and postharvest season within an apple germplasm collection on fruit softening during cold air storage. Postharvest Biology and Technology 44(3): 212-219.
Oraguzie, N.C., T. Yamamoto, J. Soejima, T. Suzuki, and N. de Silva. 2005. DNA fingerprinting of apple rootstocks using Simple sequence repeats. Plant Breeding 124: 197-202.
Oraguzie, N.C., H. Iwanami, J. Soejima, T. Harada, and A. Hall. 2004. Inheritance of the Md-ACS1 gene and its relationship to fruit softening in apple (M. x domestica Borkh). Theoretical Applied Genetics 108(8): 1526-1533.













